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Later that year, a study compared the DNA sequences using 127,000 single-nucleotide polymorphisms (mutations) of wolves and coyotes, but did not include red wolves and used Algonquin wolves as the representative eastern wolf, not wolves from the western Great Lakes states (usually referred to as Great Lakes wolves). The study indicated that Algonquin wolves were a distinct genomic cluster, even distinct from the Great Lakes states’ wolves, which it found were actually hybrids of the gray wolf and the Algonquin wolf. The study’s results did not exclude a possibility that the Great Lakes states’ wolf (the gray wolf x eastern wolf hybrid (C. l. lycaon)) historically inhabited southern Ontario, southern Quebec and the northeastern United States alongside the Algonquin wolf, as there is evidence to suggest both inhabited those areas. In 2016, a study of mDNA once again indicated the Eastern wolf as a coyote–wolf hybrid. In 2018, a study looked at the y-chromosome male lineage of canine