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Cave wolf populations are known from three caves separated by a few kilometers from each other that are located on the Franconia Karst along the Wiesent and Ahorn River vallies in the Upper Franconia region of the state of Bavaria, Germany. Sophie’s Cave sits on the northwest slope of the Ailsbach Valley near Rabenstein Castle in the Ahorntal municipality. Große Teufels Cave (Big Devil’s Cave) and the Zoolithen Cave are located nearby. They are also known from Hermann’s Cave in the village of Rübeland near the town of Wernigerode, in the district of Harz, Saxony-Anhalt, Germany. Its bone proportions are close to those of the Canadian Arctic-boreal mountain-adapted timber wolf and a little larger than those of the modern European wolf. Some postcranial bones have similarly large proportions to those from the Sophie’s and Große Teufels Caves, where the bone sizes are closer to those of Scandinavian Arctic and Canadian Columbian wolf subspecies than to those of the smaller European wolves. Bone sizes are one-eighth larger than European wolves, and this wolf was a specialized Late Pleistocene wolf ecomorph.
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Black wolf
Although the black wolves of America were originally given the same binomial name as those in Europe, Anselme Gaëtan Desmarest believed that they were a different species. Historically, the natives of the banks of the Mackenzie River, Saskatchewan River and southern Canada apparently never viewed black wolves as a distinct species. In his 1791 book Travels, William Bartram mentioned seeing black wolves among the few red wolf populations he saw in Florida. He stated that they were “perfectly black”, except the females which were described as having a white spot on the breast. Bartram also described a “black wolf-dog of the Florida Indians” which was identical to the local wolves, save for the fact that it could bark, and could be trusted around horses. The fur of a black wolf was once considered by the natives of New England to be worth over 40 beaver skins. A chieftain accepting a gift of black wolf fur was seen as an act of reconciliation. The black wolves of the Southern United States were considered a separate species to the northern kind due to differences in colour and morphology, and were named clouded or dusky wolves (Canis nubilus). The dusky wolves occurred in Missouri Territory, and were intermediate in size between common wolves and coyotes. They apparently produced a foul odour. On January 15, 2009, a black male wolf from “Mollie’s Pack” in the Yellowstone National Park’s Pelican Valley was weighed in at 143 lbs, making it the largest Yellowstone wolf on record.

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