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Modern humans’ ancestors first appeared in East Africa 195,000 years ago. Some migrated out of Africa 60,000 years ago, with one group reaching Central Asia 50,000 years ago. From there they reached Europe, with human remains dated 43,000-45,000 years BP discovered in Italy, Britain, and in the European Russian Arctic 40,000 years ago. Remains of mammoth that had been hunted by humans 45,000 YBP have been found at Yenisei Bay in the central Siberian Arctic. Another group left Central Asia and reached the Yana River, Siberia, well above the Arctic circle 27,000 years ago. Modern humans then made their way across the Bering land bridge and into North America between 20,000-11,000 years ago, after the Wisconsin glaciation had retreated but before the Bering land bridge became inundated by the sea. These people then populated the Americas. In the Fertile crescent the first agriculture was developing 11,500 years ago.

In this environment, the non-megafauna-specialist haplogroup 1 wolf became more numerous than the haplogroup 2 megafaunal wolf in the Old World and in the New World was unchallenged by the latter due to its prior extinction there.
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Japanese wolf
Before Dutch zoologist Coenraad Jacob Temminck classified it, it had been long recognized in Japan that Honshu was inhabited by two distinct canids; ōkami (wolf) and yamainu (mountain dog), both of which were described by the herbalist Ono Ranzan in his Honzō kōmoku keimō (“An instructional outline of natural studies”) in 1803. He described the ōkami as an edible, but rapacious, greyish-brown animal with a long, ash-colored, white-tipped tail with webbed toes and triangular eyes that would occasionally threaten people if rabid or hungry. In contrast, the yamainu was described as a similar animal, but with speckled yellowish fur, unwebbed toes, a foul odor and inedible meat.

Ranzan’s works were studied by German botanist Philipp Franz von Siebold during his tenure in Dejima. He purchased a female mountain dog and a wolf in 1826, describing both in his notes as distinct, and preparing two sketches illustrating their differences. The skin of the mountain dog was subsequently shipped to the Rijksmuseum van Natuurlijke Historie in the Netherlands and mounted. The specimen, along with Siebold’s notes, were used by Temminck as references for his scientific classification of the animal in Fauna Japonica (1839). Temminck, however, misinterpreted Siebold’s notes distinguishing the wolf and the mountain dog and treated the two as synonyms. In 1842, he wrote a longer description, still confounding the two names, and producing a sketch of a “wolf” based on Siebold’s mounted mountain dog specimen.

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